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As the VLP-bound MXRA8 molecules on the cell surface always adopt an “inverted” orientation with a 48-a.a.-long stalk region protruding from the C-terminal of the D1 Ig domain, a simple deletion of the stalk region in the C-terminal tagged set-up may reduce the avidity of the tag, so we replace the stalk region with a GS-linker of the same length in the C-terminal tagged set-up