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The reactions are shown here, with R and R’ representing the two gluten molecules and with cysteine as the reducing agent: R–S–S–R’ + cys–SH R–S–S–cys + R’–SH R–S–S–cys + cys–SH cys–S–S–cys + R–SH These reactions reduce the number of cross-links between the gluten subunits proportional to the number of cysteine or glutathione molecules added and are reversible so that the degree of relaxation can be controlled

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Bernfeld P & Homburger F (1983) Skin painting studies in Syrian hamsters